Οικολογία- Διαχείριση και Προστασία Φυσικού...

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ΠΑΝΕΠΙΣΤΗΜΙΟ ΠΑΤΡΩΝ ΣΧΟΛΗ ΘΕΤΙΚΩΝ ΕΠΙΣΤΗΜΩΝ ΤΜΗΜΑ ΒΙΟΛΟΓΙΑΣ ΜΕΤΑΠΤΥΧΙΑΚΗ ΔΙΠΛΩΜΑΤΙΚΗ ΕΡΓΑΣΙΑ «Οικολογία- Διαχείριση και Προστασία Φυσικού Περιβάλλοντος» Χρονισμός των αλιευτικών ενεργειών στις παραδοσιακές ιχθυοσυλληπτικές εγκαταστάσεις της λιμνοθάλασσας της Κλείσοβας με τον ημερήσιο παλιρροιακό κύκλο Καλλιόπη Σπάλα ΒΙΟΛΟΓΟΣ ΠΑΤΡΑ 2007

Transcript of Οικολογία- Διαχείριση και Προστασία Φυσικού...

-

2007

-

2007

,

: ,

2005-2007. , . , . , . , , , . , . , . , . , . . , . . , , . , , . , , , , , , . 2007,

1.

1 2 3 4 5 5 6 7 8 8 8 9 10 11 11 11 12 14 15 16 17 18 18 20 22

& - : 2.

3.

i

: Rayleigh test 4.

23 26 26 26 29 29 30 31 32 33 35 37 39 41

1: 0 cm 2: 5 cm 3: 10 cm

44 48 51 57 58 59 63 64 68 69

5.

6.

ii

7.

72 73 74 76

iii

1.

1.

1

1.

. , , . , - (Barnes 1980). : 1) , 2) , 3) , 4) , 5) , 6) 7) , / , . , , ( 2 m), , , . , (Kjerfve 1989). (Kjerfve 1989). , - , , , , ,

2

1.

(Day & Ynez-Arancibia 1982). Kjerfve (1986), , , . , . , , . . / , . . . 100 km ( km). , .

( 99%) , , , (McHugh 1967). (Day et al. 1989), . , , : 1) r- , . , / 2) - , . , , .

3

1.

/ , , , (Ynez-Arancibia et al. 1994). : 1) , 2) , , , 3) , . . . (Pauly & Ynez-Arancibia 1994). , , (Costa et al. 2002). (Katselis et al. 2003). (Kapetsky 1984). (Katselis et al. 2003).

6500 km2 (Pearce & Crivelli 1994), (Kapetsky 1984, Ardizonne et al. 1988, A 2001). , , , (Kapetsky & Lasserre 1984, Ardizonne et al. 1988, A 2001). , , ,

4

1.

.

, ( ), ( ) (Von Brandt 1984). , . (Katselis et al. 2003). (Ardizonne et al. 1988), (McDowall 1988). PLINE LANCIEN (23-79 ..). , , , , , , , , (Chaveut 1988) (Peja et al. 1996).

72 550 km2 (Kotsonias 1984). , (43,3%) (28,9%). 72% , 8% ( 2001). 89,5% ( 2001).

5

1.

147 km2, 50% . , 220 km2, ( , , ). . , , . , , . , . :A : 21 05' - 21 35 : 38 15' - 38 30'

27 km 15 km. 12 km, . , 6 ( 1) :1 2 3 4 5 6 (km2) 2,5 18 80 4,5 14 28

0,30 1,60 m, , 30 m ( et al. 1994). ( et al. 1995).

6

1.

, (Katselis et al. 2003).

1. 6 ( 1-6 ).

Ramsar, 24/1/1974 (... 350, 1 , .. 191). , , , NATURA 2000 (Sites of Community Importance - SCI) 92/43/. - , (.1650/1986, ... 477//31.5.2006).

7

1.

56 , 6 , 29 , 15 , , , 5 1 (Anguilla anguilla) ( et al. 2000).

, 16 , . , . , , . 19 , , ( 2004).

. 700 200 . , , , 10% ( et al. 1994, Katselis et al. 2003). , ( , , , , , ), .

8

1.

500 , , , , . ( et al. 1994). , , . , , , . 500 m (. 435/67). . 4,6 / km2 (Katselis et al. 2003), . (Kapetsky 1984).

, , , ( et al. 1994). , 1500-2000 tn 1960, 13001500 tn 1983 (otsonias 1984, et al. 1994), 1000-1500 tn, 45% ( et al. 2005). . 46% , 28% , 11% , 9% 6% ( et al. 1994).

9

1.

8 92,1% 1988-1998 (Katselis et al. 2003).

. , ( 2001). , , . , . [. , , . vivarium , vivus , ]. 14 , 8 6 . , . . 1826 ( & 1994). . , (Katselis et al. 2003). , . , ( et al. 1994).

10

1.

, 24, 29.5 cm ( et al. 1993). , , . ( 1996).

& - - , 25 km2. , 60, & (Hotos & Avramidou 1997).

: 19 km2. :A : 21 25 - 21 28 : 38 18 - 38 22

. , 6 km, () 1800 m 80 m. (), 1970 .

11

1.

. 3500 m 3500 m, ( 2001). 0,8 m, 1,5 m. , . (Hotos & Avramidou 1997). ( 2.600 m, 40 m 1,5 m) 1988. . 1999, , . , ( 2001).

( 2001). 1984 , . , . . ("" ""), , "(3 " "4 " "7 ). , .12

1.

"", 2007, , 30.000 . , . . 16 . , , , . . : 1. 2. 9 8 . : 70% 30% - -

: - ( )

40%

13

1.

( 2001).

, 1994-1998, 22,2 tn (Katselis et al. 2003). Mugilidae Sparidae ( Sparus aurata) (Katselis et al. 2003). 2005, 17,6 tn, (: . ).

14

2.

2.

15

2.

, . , 70% ( 2001). , , 1) , 2) , . , , . , , . , , : , , . , , . , - - .

2. .

16

3.

3.

17

3.

17/08/2005 10/02/2006, 7 . , , . , 3 , , . , 3 . , ( 3).

3. 2 . : (: Google arth), :

. , 13 mm, . 30-40 cm,

18

3.

2,5 m (1 m , 0,5 m 1 m ). 50 m. () 8 m. . 20 m. V 120 . , , . 7 (.. ). ( 4). . (), () . , .

4. 1,2,3: , 4&5: , : , : : , :

. U, 2 m 1,5 m. , .19

3.

, , 3 cm 7 cm . ( ) , , , . , , , , . ( ) , , . , . , ( , ).

, , . , , . , . , . . , . , . , . .20

3.

, . , : 1. 2. 3. 4. , , , . : 5. 6. . [ 1-4 , 5-6 ].

21

3.

2 4 , , . .

5. .

B-E (: , C: , D: & E: ). F & G .

22

3.

. . (Open University 1989). (Tsimplis 1994). T Hmax 4 : 1. .

(h)

Hmax cm 7.8 5.0 1.6 0.6

M2 12.42 92 S2 1 1 - 12.00 103 23.93 10 25.82 349

M2 t : 2 M 2 = M 2 ,max cos (t t 0 + ) 2 2 2 (t t 0 + ) = M 2 ,max cos 2 2 [1] : M 2 = 7, 8 cos 0, 1610 ( t + 1, 606 ) (M2 cm, t h)

[1]

[2].

S2, 1, 1 : S2 = 5, 0 cos 0, 1666 ( t + 1,798 ) (S2 cm, t h) [3] K 1 = 1, 6 cos[0, 0835 (t + 0, 174)] (K1 cm, t h) O1 = 0, 6 cos[0, 0774 (t + 6, 091)] (O1 cm, t h)

[4] [5]

[2], [3], [4] [5] ht t.23

3.

ht = 7, 8 cos 0, 1610 ( t + 1, 606 ) + 5, 0 cos 0, 1666 ( t + 1,798 ) +1, 6 cos 0, 0835 ( t + 0, 174 ) + 0, 6 cos 0, 0774 ( t + 6, 091 ) , 6/10 - 21/11/1998. , . (smoothing analysis), variability 48 h. (N: 3820, E: 2125) (N: 3814, E: 2144) t0 = 3 , 5h . 1/10/2002. , , , GMT, GMT. 1 hmaxj j f = TM 2 2 . f = 0, 08 h-1. , - 0 ( ). j ( ) t i. : h t = h max j cos i . [7] = Ti : 2 2 = 12,42 h, [7] 2

24

3.

i =

ht 6.21 cos1 ( h) h max j i

[8]

i 8 0 ( ) 6.21 ( ). i > 6.21 h 6.21 .

25

3.

, - , , . , , , 2 .

a , X [9] cos a = r

sin a =

r

[10]

[11] n n [9] [10] Y tan a = [12]. X [12] a rad , Ti ( ) (Zar 1999).

:

X=

cos ai =1

n

i

,Y=

sin ai =1

n

i

r = X 2 + Y 2

r, . 0 1. r=0, , r=1 . r

26

3.

, 1- r . 1 r = 1 , r=0 ( 6), , 1 r = 0 r=1 ( 7).

6. . r, .

7. . r, .

r=0 , ( 8).

8. . r, .

27

3.

(angular variance) : s 2 = 2 (1 r ) ( rad 2 ) [13]

0 2, . [12] : 6, 21 2 (1 r ) ( h2) s2 = [14].

(angular deviation) :

s = 2 (1 r ) ( rad)

[15]

[15] : 6, 21 2 (1 r ) ( h) s= [16].

[16] : s ( h) [17] sT = n (Zar 1999).

d . r 0, 92 r > ,1 /2 n2 2 n(2 R 2 n ,1 2 4n ,1 d = arc cos R

( rad)

[18]

2 2 n(2 R 2 n ,1 2 4n ,1 6, 21 d= arc cos R

( h)

[19],

R = nr . 2 ,1 =0,05, =1 ( 95%) 3,841 (Zar 1999).

28

3.

, . r 0 ( ). , ( 8). . 2 ai , r [11] , : X cos 2 a = [20] sin a = [21] r r Y [22] [22] [20] [21] tan 2 a = X 2a ( rad). 2a =a. 2 s [15], ( rad) 2 [16]. r 0, (Zar 1999).

: Rayleigh test , a , , r. ( ), H0 : z Rayleigh. R2 [23]. z= = nr 2 n

29

3.

z > z ,n H0 , , z < z ,n , (Zar 1999).

, , . Watson-Williams, 0: T1 = T2 = .... = Tk .

F : k (N k ) R j R j =1 F= [21]. k ( k 1) N R j j =1 k , R R N = j =1 n j .n

F F(1), k 1, N k =0.05 (Zar 1999).

30

4.

4.

31

4.

9. .

9 , ( 5 cm). 3 : 1. 0 cm 2. 5 cm 3. 10 cm 2. 3 .

hmaxj

0-4 (cm)

5-9 (cm) 10 (cm) % % % 100,00-81,82 100,00-84,35 100,00-88,07 100,00-93,24 72,73-35,23 76,52-38,26 81,65-42,20 86,49-48,65 29,55 27,83 33,94 40,54

32

4.

100,00-87,94 100,00-91,07

82,98-51,77 86,52-44,94

43,97 39,33

2 , 80% ( ) hmaxj 5 cm, 80% 40% , hmaxj ( 9 cm). , hmaxj 10 cm 30% .

1: 0 cm 0 cm

, , . ( 10, 1). . , , ( 10, 2).

10. (0) hmaxj 0 cm.

33

4.

0 cm

, ( 11, 1). , . , ( 11, 2). . , ( 11, 3). , , , ( 11, 4).

11. (0) hmaxj 0 cm.

34

4.

2: 5 cm 5 cm

. , , , ( 12, 1). . 2 , , . ( 12, 2).

12. (0) hmaxj 5 cm.

35

4.

5 cm

. , , ( 13, 1). , ( 13, 2). , . , ( 13, 3). , ( 13, 4).

13. (0) hmaxj 5 cm.

36

4.

3: 10 cm 10 cm

, , ( 14, 1). ( 14, 2).

14. (0) hmaxj 10 cm.

37

4.

E 10 cm

. , , ( 15, 1). ( 15, 2). , ( 15, 3). . , ( 15, 4).

15. (0) hmaxj 10 cm.

38

4.

3: ( T ), (r), (n), ( sT ) 95% (d) 3 .

0 cm

5 cm

10 cm d () 0.46 0.70 0.82 -

T () 7.38 0.95 10.70 2.39 11.77 5.40

r 0.5589

n 141

sT() 0.16 0.21 0.20 0.29 0.25 0.30

d () 0.38 0.58 0.59 -

T ()7.56 1.16 10.77 5.24 8.99 5.81

r 0.5144 0.4345 0.3492 0.0731* 0.0118* 0.1678*

n 117 77 91 66 90 64

sT() 0.18 0.24 0.24 0.33 0.29 0.32

T ()7.90 2.21 11.49 5.38 4.65 4.82

r 0.5382

n 62

sT() 0.24 0.31 0.40 0.50 0.38 0.38

d () 0.61 0.75 1.49 1.64 1.18

0.4807 89 0.4193 118 0.0447* 90 0.1028* 111 0,1261* 74

0.5829 35 0.3522 32 0.1709* 26 0.3058 0.4352 37 30

(*) r, T . , . 4: Rayleigh test H0

z 44.046 20.562 20.745 0.180 1.173 1.176

0 cm

5 cm H0 z

10 cm z

z0.05,n2.990 2.987 2.990 2.987 2.989 2.986

z0.05,n2.990 2.986 2.987 2.984 2.987 2.984

H0

z0.05,n2.983 2.975 2.972 2.967 2.975 2.971

H0 39

30.957 14.536 11.095 0.353 0.012 1.802

17.958 11.892 3.970 0.759 3.461 5.682

4. 5: ( T ), (r), (n), ( sT ) 3 . .

0 cm

T () 5.34/12.16 2.71/8.92 5.95/11.55

r

n

sT ()0.14 0.13 0.15

0.1297 90 0.0828 111 0.1054 74

5 cm r n sT () T () 4.25/10.46 0.1292 66 0.16 3.10/9.31 0.1574 90 0.14 5.86/12.07 0.1341 64 0.16

10 cm

T ()3.59/9.80 -

r

n

sT ()0.25 -

0.1497 26 -

3 4 11, 13 15 , 0 5 cm, 10 cm .

40

4.

(0). , 3 : 1. : 0 cm 1 2 3 4 5 6

T sT (h)7.38 0.16 10.70 0.20 0.95 0.21 2.71 0.13* 8.92 0.13* 5.34 0.14* 12.16 0.14* 5.95 0.15* 11.55 0.15*

2. : 5 cm

T sT (h)

7.56 0.18 1 2 10.77 0.24 3 1.16 0.24 3.10 0.14* 4 12.07 0.14* 4.25 0.16* 5 10.46 0.16* 5.86 0.16* 6 12.07 0.16*

3. : 10 cm 1 2 3 4 5 6

T sT (h)

7,90 0.24 11.49 0.40 2.21 0.31 3.59 0.25* 9.80 0.25* 4.65 0.38 4.82 0.38

sT ( ) .41

4.

, 3 , . , . 3 , , 3,5 , . . . . 3 . 3 . , . , , . , 0 5 cm , . . 10 cm, . , , , . 3 . , ,

42

4.

, , . , , ( r sT ) ( r sT ). 3 . . , 0 5 cm, . ( ) ( r). , sT , , ( ) . . , , . , ( r) 3 . , sT .

43

4.

6: ( T ), (r), (n), ( sT ) 95% (d) 3 .

0 cm

5 cm d () () 0.43 1.05 0.34 1.0 0.29 0.68 0.44 1.47 0.33 0.77 0.46 1.16 0.77 -

10 cm

T ()7.61 7.29 7.41 7.95 7.45 6.51 7.32

r 0.7365 0.4368 0.6498 0.4089 0.7363 0.6200 0.8465

n 11 39 32 24 19 14 2

sT

T ()7.75 7.37 7.64 8.68 7.45 6.67 7.32

r 0.7085 0.3393 0.6087 0.3921 0.7363 0.6088 0.8465

n 9 31 23 20 19 13 2

sT() 0.50 0.41 0.36 0.49 0.33 0.48 0.77

d () 1.26 1.60 0.90 1.76 0.77 1.25 -

T ()7.36 8.61 8.24 9.49 7.02 6.16 8.43

r

n

sT() 0.60 0.54 0.49 0.51 0.46 0.53 0.00

d () 1.64 1.80 1.22 1.32 1.14 1.37 -

0.6735 7 0.4437 15 0.6946 10 0.6311 11 0.7552 9 0.6737 9 1.0000 1

7: ( T ), (r), (n), ( sT ) 95% (d) 3 .

0 cm

5 cm

10 cm

T ()1.37 1.33 1.24 0.53 11.72 1.01

r 0.0000 0.4958 0.5373 0.5312 0.4156 0.6208 0.6239

n 0 24 20 15 19 9 2

sT() 0.00 0.41 0.43 0.49 0.49 0.57 1.21

d () 1.14 1.14 1.39 1.68 1.55 -

T ()1.84 1.67 1.23 1.03 11.72 1.01

r 0.0000 0.4070 0.5077 0.4674 0.4255 0.6208 0.6239

n 0 19 17 13 17 9 2

sT() 0.00 0.49 0.48 0.57 0.51 0.57 1.21

d () 1.73 1.36 1.85 1.75 1.55 -

T ()2.69 2.95 2.13 2.19 11.49 2.79

r

n

sT() 0.00 0.51 0.50 0.72 0.64 0.60 0.00

d () 1.28 1.27 2.22 1.81 1.67 44

0.0000 0 0.6721 10 0.7437 8 0.6725 5 0.6824 6 0.7732 5 1.000 1

4.

8: ( T ), (r), (n), ( sT ) 95% (d) 3 .

0 cm

5 cm

10 cm

T ()10.62 11.00 12.25 9.52 8.99 8.91 -

r 0.5529 0.3780 0.5325 0.4134 0.9478 0.8869 0.0000

n 17 48 27 12 3 11 0

sT() 0.45 0.32 0.37 0.62 0.37 0.28 -

d () 1.21 1.07 0.97 2.58 1.71 0.78 -

T ()10.82 11.14 0.55 8.55 8.71 8.84 -

r 0.5716 0.3072 0.5976 0.4400 0.9493 0.8810 0.0000

n 14 37 19 9 2 10 0

sT() 0.49 0.38 0.41 0.70 0.45 0.30 -

d () 1.31 1.63 1.02 0.83 -

T ()10.99 10.35 1.72 0.26 9.31 -

r

n

sT() 0.72 0.62 0.35 0.18 0.74 -

d () 1.19 2.69 -

0.4019 9 0.3072 13 0.9231 5 0.9913 2 0 0.7869 3 0.0000 0

9: ( T ), (r), (n), ( sT ) 95% (d) 3

0 cm

5 cm

10 cm

T ()9.38 0.89 5.02 4.07 11.22 8.72 -

r

n

sT() 0.91 0.39 0.52 0.54 0.66 0.64 -

d () 1.43 2.43 2.03 -

T ()8.38 1.00 5.04 4.28 10.45 8.36 -

r

n

sT() 0.32 0.45 0.58 0.37 0.15 0.59 -

d () 1.11 2.64 1.59 -

T ()8.38 0.47 4.34 4.29 8.98 -

r

n

sT() 0.32 0.73 0.55 0.46 1.08 -

d () 1.68 1.84 -

0.4736 5 0.2158 40 0.2175 23 0.6240 10 0.8320 3 0.5300 9 0.0000 0

0.9736 2 0.1879 32 0.3223 16 0.8964 6 0.9939 2 0.6443 8 0.0000 0

0.9736 2 0.0478 14 0.8478 4 0.9191 3 0 0.5542 3 0.0000 0

45

4.

10: ( T ), (r), (n), ( sT ) 95% (d) 3 .

0 cm

5 cm

10 cm

T ()12.09 11.53 1.78 9.62 12.36 8.05 6.82

r 0.9255 0.1200 0.3811 0.1767 0.1288 0.4502 0.6827

n 6 34 26 16 11 13 5

sT() 0.31 0.45 0.43 0.63 0.79 0.57 0.70

d () 1.04 1.53 1.97 2.14

T ()11.68 11.52 2.59 8.38 1.48 7.98 6.82

r 0.9714 0.1200 0.4149 0.1289 0.1088 0.6005 0.6827

n 4 27 20 13 10 11 5

sT() 0.24 0.50 0.48 0.72 0.83 0.53 0.70

d () 1.26 1.62 1.42 2.15

T ()11.87 5.09 3.56 4.91 4.24 6.84 5.08

r

n

sT() 0.22 0.76 0.47 0.55 0.94 0.67 0.00

d () 1.57 1.28 1.68 1.95 -

0.9810 7 0.1200 15 0.8293 10 0.8466 11 0.4330 9 0.6560 6 1.0000 1

11: ( T ), (r), (n), ( sT ) 95% (d) 3 .

0 cm

5 cm

10 cm d () -

T ()7.90 2.16 5.61 4.73 10.72 7.30 7.12

r

n

sT() 0.76 0.57 0.58 0.60 0.92 0.92 -

d () -

T ()7.90 2.67 5.30 5.39 8.80 8.12 7.12

r

n

sT() 0.76 0.60 0.62 0.64 0.97 0.89 0.99

T ()7.90 3.52 4.25 4.91 4.39 6.48 3.49

r 0.7770 0.4290 0.8648 0.3495 0.6465 0.9908 0.4924

n 3 7 5 9 2 2 2

sT() 0.76 0.80 0.46 0.75 1.18 0.19 1.41

d () 1.35 2.70 -

0.7770 3 0.3006 17 0.2293 18 0.2129 17 0.1323 8 0.3488 6 0.3768 5

0.7770 3 0.3200 15 0.3087 14 0.2030 15 0.1549 7 0.4922 5 0.3768 5

46

4.

12: Watson-Williams 3 .

P-value 0.8235 0.5700 0.0067 0.0000 0.0002 0.0476

0 cm F Df, (n) 0.4784 7, (141) 0.8035 7, (89) 3.1631 7, (118) 5.8282 7, (90) 4.9345 7, (114) 2.26308 6, (74)

0 NS NS S S S S

5 cm P-value F Df, (n) 0 0.6330 0.7217 7, (117) NS 0.5145 0.8796 7, (77) NS 0,0010 4.1793 7, (91) S 0,0003 5.0692 7, (66) S 0,0001 5.5577 7, (90) S 0,0834 1.9810 7, (64) NS

P-value 0.0469 0.0798 0.1521 0.3407 0.0443 0.3476

10 cm F Df, (n) 2.3042 7, (62) 2.1414 7, (35) 1.744 7, (32) 1.2169 7, (26) 2.4977 7, (37) 1.1873 7, (30)

0 S NS NS NS NS NS

47

4.

3 , . 3 , , . . , , . . 0 5cm 10 cm ( 12). , 10 cm, : -- , -- , , . 16. , 3 . 0 5 cm, , ( 12). , , . . 10 cm, , 2 , . 17.

48

4.

16. (0) ( 2* sT ).

49

4.

17. (0) ( 2* sT ). 50

4.

. , 3 . , ( ) . , 0 5 cm ( 12). , , , , . 10 cm . 18. . 0 5 cm, 2 ( 12). , , , . 19.

51

4.

18. (0) ( 2* sT ). 52

4.

19. (0) ( 2* sT ). 53

4.

0 5 cm, 10 cm. , 2 ( 12). , 0 5 cm, , . , . , , , . T 20. 0 5 cm 0 cm. 10 cm ( 12). , 0 5 cm , , , . 10 cm, , - , . 21.

54

4.

20. (0) ( 2* sT ). 55

4.

21. (0) ( 2* sT ). 56

5.

5.

57

5.

3 . , . , , . , , . , , . , . , , . , . . , , , . , . , () () , . , .

58

5.

, , , , . , .

, , , , . , , . . , . . . , , . , , . . , . 3 , . 3.5 . , 59

5.

3 . , , . , , , . , , 10 cm, , . , . , , : - , - . , 2 (0 5 cm), , (10 cm) . 0 5 cm, , . , , , . : 1) ,

60

5.

( , 2) , , ( ) , , . . . , . , . . , , . , , . , , ( ) . 10 cm, . . , , , . . , 3

61

5.

. , . . , , , . , , , . , , .

62

5.

, . , . 3 , , , , . , , . , . . , . , . , , 10 cm , . 3 , . , .. .

63

5.

. . , . , . Mugil cephalus (Katselis et al. 2003). , , 140 euro/kg (Katselis et al. 2005). , . , . , , . , . (Katselis et al. 2003). . , , Sparus aurata. , . . , . , , .

64

5.

, . . , . , Gobius sp., , . , . 0 5 cm. , . . , , ( ). . . , , . 10 cm, , . , 1 6 . . , .

65

5.

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., ., . &. . (1995). . 4 . 27-41. (2001). & . / & . . . ., ., . & . (1994). . 155: 82-91. ., ., . & . (2005). Lisa saliens () . 12 , , 2005. . (1996). (Aphanius fasciatus Nardo, 1827) . . 198 . ., . & . (2000). . 6 2: 239-244. . (2004). . (18 2 ). , 193 . ., ., . & (1993). - . . . ..., , & ACE-2242/87/10-1, , , . . & . (1994). . . , 1994.

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