ISSN 1020-6868

FAO SPECIES IDENTIFICATION GUIDE FOR FISHERY PURPOSES

FIELD IDENTIFICATION GUIDE TO THE LIVINGMARINE RESOURCES OF THE EASTERN ANDSOUTHERN MEDITERRANEAN

FAO SPECIES IDENTIFICATION GUIDE FOR FISHERY PURPOSES

FIELD IDENTIFICATION GUIDE TO THE LIVINGMARINE RESOURCES OF THE EASTERN ANDSOUTHERN MEDITERRANEAN

by

M. Bariche

Department of Biology, American University of Beirut, Lebanon

FOOD AND AGRICULTURE ORGANIZATION OF THE UNITED NATIONS

Rome, 2012

2012Index

The designations employed and the presentation of material in thisLQIRUPDWLRQSURGXFWGRQRWLPSO\WKHH[SUHVVLRQRIDQ\RSLQLRQZKDWVRHYHUon the part of the Food and Agriculture Organization of the United1DWLRQV)$2FRQFHUQLQJWKHOHJDORUGHYHORSPHQWVWDWXVRIDQ\FRXQWU\territory, city or area or of its authorities, or concerning the delimitationRILWVIURQWLHUVRUERXQGDULHV7KHPHQWLRQRIVSHFLFFRPSDQLHVRUSURGXFWVRIPDQXIDFWXUHUVZKHWKHURUQRWWKHVHKDYHEHHQSDWHQWHGdoes not imply that these have been endorsed or recommended by FAOin preference to others of a similar nature that are not mentioned.7KHYLHZVH[SUHVVHGLQWKLVLQIRUPDWLRQSURGXFWDUHWKRVHRIWKHDXWKRUVDQGGRQRWQHFHVVDULO\UHHFWWKHYLHZVRI)$2

ISBN 978-92-5-006450-5

All rights reserved. FAO encourages reproduction and disseminationRIPDWHULDOLQWKLVLQIRUPDWLRQSURGXFW1RQFRPPHUFLDOXVHVZLOOEHauthorized free of charge, upon request. Reproduction for resale or othercommercial purposes, including educational purposes, may incur fees.Applications for permission to reproduce or disseminate FAO copyrightmaterials, and all queries concerning rights and licences, should beaddressed by e-mail to copyright@fao.org or to the Chief, Publishing3ROLF\DQG6XSSRUW%UDQFK2IFHRI.QRZOHGJH([FKDQJH5HVHDUFKDQG([WHQVLRQ)$29LDOHGHOOH7HUPHGL&DUDFDOOD5RPH,WDO\ FAO 2012

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PREPARATION OF THIS DOCUMENT

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Fischer et al 6HUHQD &,(60 $WODV RI ([RWLF 6SHFLHV LQ WKH0HGLWHUUDQHDQ )$2 )LVKHULHV DQG $TXDFXOWXUH ,QIRUPDWLRQ DQG 6WDWLVWLFV6HUYLFH

Index

iii

ivBariche, M.)LHOGLGHQWLFDWLRQJXLGHWRWKHOLYLQJPDULQHUHVRXUFHVRIWKH(DVWHUQDQG6RXWKHUQ0HGLWHUUDQHDQ)$26SHFLHV,GHQWLFDWLRQ*XLGHIRU)LVKHU\3XUSRVHVRome, FAO. 2012. 610 pp.

610 .2012

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372

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Abstract

Acknowledgements

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vi,QWHUXQLYHUVLWDULR GL %LRORJLD 0DULQD HG (FRORJLD$SSOLFDWD /LYRUQR ,WDO\ IRUFDUWLODJLQRXVVKHVDQG*LXOLD0R,VWLWXWR6XSHULRUHSHUOD3URWH]LRQHHOD5LFHUFD$PELHQWDOH5RPH,WDO\IRUSLQQLSHGVSincere thanks go to the FAO team for providing full assistance at all levelsGXULQJWKHSUHSDUDWLRQRIWKHPDQXVFULSWIURPGDWDJDWKHULQJWRQDOHGLWLQJ7KDQNVWR1LFROHWWD'H$QJHOLV(PDQXHOD'$QWRQL0LFKqOH.DXWHQEHUJHULongo and Monica Barone for their patience and dedication. I am also gratefulto Piero Mannini and Michel Lambuf for giving me the opportunity to produceWKLV ZRUN WR ,VVDP .URXPD )$2 &RQVXOWDQW 6\ULD IRU WKH WUDQVODWLRQ LQWR$UDELFDQGIRU$UDELFWHFKQLFDOHGLWRULDOZRUNDQGWR6DPLU0DMGDODQL)LVKHULHVDQG:LOGOLIH0LQLVWU\RI$JULFXOWXUH/HEDQRQIRUKLVFRQWLQXRXVVXSSRUWDQGhelp. I am also indebted to Mohamad Abdul Sater, Hana Ibrahim, Paul Ramia,Colin Smith and Nancy Sayar for their assistance and valuable help at differentOHYHOVRIWKLVZRUN,DPDOVRJUDWHIXOWR7RXIHHN$VVDOSURIHVVLRQDOVKHUPDQIURP%DWURXQIRUhis friendship, constant help and availability and for his incredible love to the0HGLWHUUDQHDQ6HD7KDQNVDUHDOVRH[SUHVVHGWRDOOWKH/HEDQHVHVKHUPHQIRU SURYLGLQJ PH ZLWK YDOXDEOH LQIRUPDWLRQ VSHFLPHQV DQG IRU UHJXODUO\reporting their catches.7KLV ZRUN LV GHGLFDWHG WR DOO VKHUPHQ DQJOHUV DQG SDVVLRQDWH RI WKHMediterranean Sea.

Table of Contents

PageINTRODUCTION . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .1

1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

A PICTORIAL INDEX TO DIVISIONS AND FAMILIES . . . . . . . . . . . . . . . 40

40 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

GLOSSARY . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 52

52 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

INTRODUCTION TO ARTHROPODS . . . . . . . . . . . . . . . . . . . . . . . . 68

68 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

STOMATOPODS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 68Order Stomatopoda Mantis Shrimps . . . . . . . . . . . . . . . . . . . . 69SQUILLIDAE . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 69

68 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 69 . . . . . . . . . . . . . . . . . . . . . . . . . . .

69 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

SHRIMPS AND PRAWNS . . . . . . . . . . . . .TECHNICAL TERMS AND MEASUREMENTS ,QIUDRUGHU3(1$(,'($3HQDHLG6KULPSV .SOLENOCERIDAE . . . . . . . . . . .ARISTEIDAE . . . . . . . . . . . . . .PENAEIDAE . . . . . . . . . . . . . .SICYONIIDAE . . . . . . . . . . . . . ,QIUDRUGHU&$5,'($&DULGHDQVKULPSV . .PASIPHAEIDAE . . . . . . . . . . . .PALAEMONIDAE . . . . . . . . . . .PANDALIDAE . . . . . . . . . . . . .

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LOBSTERS . . . . . . . . . . . . . . . . . . . . .TECHNICAL TERMS AND MEASUREMENTSNEPHROPIDAE . . . . . . . . . . . .PALINURIDAE . . . . . . . . . . . . .SCYLLARIDAE. . . . . . . . . . . . .

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CRABS. . . . . . . . . . . . . . . . . . . . .TECHNICAL TERMS AND MEASUREMENTS ,QIUDRUGHU%5$&+1 000 m depth)(after Millot (1987), Lacomb et al. (1985) for the western basin. ThePRYHPHQWVRIWKHGHHSFXUUHQWVLQWKHHDVWHUQEDVLQKDYHEHHQVLPSOLHG(Roether and Schlitzer, 1991; Millot and Taupier-Letage, 2004)

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