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, " ," ().

. , 100 . ( ). ,

- . ( ), .

" - .

( 15 ) ; , , ; , .

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, Michigan . . , .

O COMPUTER

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. , 100 .

.

on / off . (off) (n).

, ( ""

), . (Synapse) . (). , axons

? 60watt. . ().

"" . .

,, . , ; , . , . .

, , .

-REM ( )

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-.

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S

( ) ON OFF ( ); ,

,

. . , . , , , . . , .

. , , . "off" . . ,

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.

-REM , . von-Economo, ,1,2 ,

, . , 3 .4

. .5 Fos,

,6

Fos .7 , Fos - , .

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-----------------------------------------------------

1 - . . ( -REM), REM.

EEG= , EMG = ( Szymusiak

,15

).

-----------------------------------------------------

GABA-. 8, 9 10-13 , . NREM ,

, .14, 15 , , D2, 1 [growth hormone releasinghormone]. 16 , , NREM, ,

. - o REM. ,
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, NREM . - NREM REM, NREM REM.8, 16

NREM. , , .

()

[excitatory/inhibitory postsynapticpotentials: EPSPs/IPSPs] . [sleep spindles] . Steriade,McCormick 17-22 , , .

GABA- . 7 14 Hz, , , . , , . GABA , . IPSPs

, . , EPSPs IPSPs , . , .

, ,

. , NREM, REM.

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[locus coeruleus], [raphe nuclei] , 23-26 .16 ( GABA- )

NREM . , GABA, GABA- , .16, 27-29 , , . , , ,

.30 , , [ascending activating system], , .31-35

REM

REM

. 2 REM. , . REM. 24 REM ( ), (). . 36 (.. ) ,

.37 , .38 REM [tonic] .

.39 , , -- [ponto-

geniculooccipital

(PGO) spikes]. , 30 REM .

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REM, 3 10 , . -- , REM. , , , . ,

, .40 , REM. REM, REM, . . ,

PGO . , .41-43 REM [phasic] .

-----------------------------------------------------

2 : - . : 4 - . REM = , G =- , EOG =, OLF= , LGN= , HIPP= , EMG= , PGO=-- .

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-----------------------------------------------------

3 L= 1,6 Berman [BermanAtlas] . RN

= , LC = , 6 = , 7= , = . H() - () .

-----------------------------------------------------

4 . . EMG = , EKG
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= , RESP = . = 50 mV ( Siegel 49)

-----------------------------------------------------

, REM ,

.47 , REM. ( 3, ). , .48 , REM- REM,

, REM ( 4).49 , , , REM , REM.

, REM ( 2 , 5). 50 PGO PGO REM REM .

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-----------------------------------------------------

5 : (), () (LGN) - . 1 . ,

. ( Siegel.51)

-----------------------------------------------------

, PGO, REM. 49, 51

, , ,

REM, , , REM. REM , REM. Matsuzaki . 52 ,

, PGO , REM .

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REM. REM. , PGO , ( 6, ). 63 PGO

REM.

. REM , REM , PGO, . , , ( 6,). 64, 65 REM ,

, . , . REM . 53,66-69

.

Sylvious [periaqueductal gray] ( - ) REM, REM . , GABAA REM , .70 71 .35 ,

[deep mesencephalic nucleus].

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-----------------------------------------------------

6 - REM , . () REM ( ). , , -- (PGO) REM. , REM

PGO REM. ( Shouse& Siegel,63 Elsevier Science.)

-----------------------------------------------------

GABA ( [pontine oralis] ) REM GABA , .72, 73

, GABA, GABA

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. GABA REM .74

REM, George . 75

REM . REM REM, REM . . , , REM

. 76-78 PGO, PGO , , . , REM ,79-81 , .82

REM . . 31, 48, 83 , .31, 84-86

REM .34, 87 , REM,88 , .

,

, REM , .

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, . [unit recording] REM.

, REM REM. 41, 42, 58, 89, 90 REM , REM. REM . ,

, , , . , . , REM, REM , REM. REM

,59, 91 92 .

REM. [microdialysis studies] REM, REM, .93 .

REM (REM- [REM-on cells]). , REM.94 , REM- . 95

REM 96, .35 Fos [tract tracing]

REM. 31, 34, 35, 48, 8486, 97

---------------------------------------------------

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7 REM- [REM sleep-off] . EEG = , EOG=, LGN= , EMG =.

-------------------------------------------------

. , 98,98 100 , 25 , NREM, REM ( 7). , REM, REM. REM GABA , 27-29, 101 REM- GABA- . 102, 103 GABA REM, 28 REM.

PGO. 104, 105 , . PGO .106-108 ,
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NREM REM PGO REM.

REM Fos 20 . Quattrochi

PGO - . . 108-110

Fos REM . Verret 111 -

Fos REM. , Fos - , -, Sylvious . , REM. and , [paragigantocellular] 112

[raphe obscurus]. , GABA- REM.29 Verret 83 Sylvious , GABA- , REM. Maloney GABA- Fos REM.

REM , Sylvious GABA- GABA- , 28, 113 .

Fos , REM.

, REM Fos REM, REM ,

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NREM .8 Fos [melanin concentrating hormone neurons] , Fos REM REM .114, 115

REM .

. NREM . REM , .116

NREM REM . REM REM. REM .

REM

. 116, 117 GABA, .118 , . REM [magnocellularis] -

[paramedianus] ,117 . 31

[ambiguous] . REM .

,

. 119 , GABA , ,

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, REM [disfacilitation].

[facilitatory] . [pontine inhibitory area, ] , 31 .

, . 120

. REM (REM- ) , , ( 8).

GABA REM .85, 86 , ( 9). 31, 84 , , ( 9). 48, 121

. 122, 123

, . , - (

REM) - , . 122

[jawappliances] . ,

[bruxism]. ,

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.124 , , NREM. , REM. , GABA NREM,

125 REM.126 , REM. NREM. REM. , REM, --D-

(NMDA) -NMDA REM. 127

, , REM. REM

. 128 , - , . (. ). 24 ,

, , .

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9. .

. . (

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74_ΕΓΚΕΦΑΛΟΣ ΚΑΙ ΑΝΤΙΒΑΡΥΤΗΤΑ_GR

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